Bythotrephes longimanus

Common Name: Spiny waterflea

Synonyms and Other Names:

Formerly known as Bythotrephes cederstroemii (Berg and Garton, 1994; Yan and Pawson, 1998; Berg et al., 2002; Therriault et al., 2002).  spiny water-flea



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Identification: Bythotrephes longimanus is a large cladoceran distinguished by a long straight tail spine that is twice as long as its body and has one to three pairs of barbs. Parthenogenically produced animals have kink in middle of their spine and sexually produced animals lack the kink. Bythotrephes appearance is similar to Cercopagis pengoi, another Great Lakes invader, except Bythotrephes is larger with a more robust spine that lacks a hook at the end.


Size: 6mm to 15mm


Native Range: Northern Europe and Asia


Great Lakes Nonindigenous Occurrences: Bythotrephes was first detected in December 1984 in Lake Huron (Bur et al., 1986), then Lake Ontario in September 1985 (Lange and Cap, 1986), Lake Erie in October 1985 (Bur et al., 1986), Lake Michigan in September 1986 (Evans, 1988), and Lake Superior in August 1987 (Cullis and Johnson, 1988).

 


Table 1. Great Lakes region nonindigenous occurrences, the earliest and latest observations in each state/province, and the tally and names of HUCs with observations†. Names and dates are hyperlinked to their relevant specimen records. The list of references for all nonindigenous occurrences of Bythotrephes longimanus are found here.

Full list of USGS occurrences

State/ProvinceFirst ObservedLast ObservedTotal HUCs with observations†HUCs with observations†
IL198620112Lake Michigan; Little Calumet-Galien
IN198619881Lake Michigan
MI1984202012Betsie-Platte; Betsy-Chocolay; Dead-Kelsey; Keweenaw Peninsula; Lake Huron; Lake Michigan; Lake Superior; Michigamme; Ontonagon; Pere Marquette-White; St. Marys; Thunder Bay
MN198720224Baptism-Brule; Cloquet; Lake Superior; St. Louis
NY198520226Lake Champlain; Lake Erie; Lake Ontario; Mettawee River; Oneida; Seneca
OH198520191Lake Erie
ONT20012003*
PA199820021Lake Erie
VT201420251Lake Champlain
WI198620188Bad-Montreal; Beartrap-Nemadji; Door-Kewaunee; Lake Michigan; Lake Superior; Lower Fox; Menominee; St. Louis

Table last updated 4/14/2026

† Populations may not be currently present.

* HUCs are not listed for areas where the observation(s) cannot be approximated to a HUC (e.g. state centroids or Canadian provinces).


Ecology:  

HABITAT: Bythotrephes longimanus is found among the zooplankton in the upper water column of large and small temperate lakes. They can tolerate brackish water, and are most abundant in late summer and autumn. Temperature plays a major role in determining the abundance and location of Bythotrephes in the Great Lakes, as they prefer cooler water and cannot tolerate very warm lake temperatures (Berg and Garton 1988; Garton et al. 1990; Brown and Branstrator 2004). Bythotrephes occurs in oligotrophic and mesotrophic lakes and has a lower tolerance to low dissolved oxygen concentrations than the native cladoceran Leptodora kindtii (Sorensen and Branstrator 2017).

 

Environmental Parameter Value Source
Thermal tolerance 4 and 30°C Grigorovich et al. 1998
Thermal preference  10 and 24°C Grigorovich et al. 1998
Salinity tolerance 0.04 and 8.0 ppt Grigorovich et al. 1998
Salinity preference 0.04 and 0.4 ppt Grigorovich et al. 1998

 

FOOD WEB: Bythotrephes longimanus is a visual predator, using its large compound eye to detect zooplankton (Azan et al. 2015). They consume 75% of their body weight each day in prey items (Lehman et al. 1997).

LIFE HISTORY: Bythotrephes can reproduce both asexually and sexually; unfertilized eggs are carried in a brood pouch, and fertilized eggs are cast in the fall, hatching the following spring (Evans 1988). The intensity and type of predation pressure appears to affect the size of Bythotrephes, its spine length, and the extent of its diel migrations (Straile and Halbich 2000).


Means of Introduction: Bythotrephes was probably introduced from ship ballast water (Sprules et al. 1990, Berg et al. 2002) and possibly as diapausing eggs from sediment in ballast tanks (Evans 1988). 


Great Lakes Status: Widespread, overwintering and reproducing.


Great Lakes Impacts:
Summary of species impacts derived from literature review. Click on an icon to find out more...

EnvironmentalSocioeconomicBeneficial



Bythotrephes longimanus has a high environmental impact in the Great Lakes.

Bythotrephes reproduce rapidly and consume small zooplankton such as small cladocerans, copepods, and rotifers, potentially competing directly with planktivorous larval fish for food (Berg and Garton, 1988; Evans, 1988; USEPA, 2008; Vanderploeg et al., 1993). At times, its consumption may even exceed that of zooplankton production (Bunnell et al., 2011). Bythotrephes, in combination with zebra mussels, is associated with slower walleye growth. Following invasion by Bythotrephes or zebra mussels, walleye were on average 12% and 14% smaller, respectively, at the end of their first summer, likely due to changes in zooplankton communities (Hansen et al. 2020). It is important to note that predation by Bythotrephes on rotifer predators and competitors has also been shown to reduce both mortality and competition, leading to food web alterations and positive effects on rotifer populations, particularly Conochilus (Pothoven and Vanderploeg 2022; Marshall et al. 2024).

A decline in native cladocerans following the introduction of Bythotrephes has been observed in Lake Huron and Lake Michigan (Barbiero and Tuchman, 2004). In Lake Erie, the detection of Bythotrephes in 1985 was also accompanied by a decline in multiple species of cladocerans (e.g., Eubosmina coregoni, Daphnia mendotae, D. retrocurva), including an almost complete absence of Diaphanosoma spp. wherever Bythotrephes was present in 1986 (Barbiero and Rockwell, 2008). This study further documented a lack of spatial overlap between Bythotrephes and Leptodora (a native water flea). This is in concordance with many other studies which have documented a negative correlation between the abundance of the two species, implying that competition and/or predation from Bythotrephes has played a significant role in declines of Leptodora (e.g., Branstrator, 1995; Fernandez et al., 2009; Foster and Sprules, 2009; Garton et al., 1990; Lehman and Cáceres, 1993; Yan and Pawson, 1997). Interestingly, Leptodora coexists with and often outnumbers Bythotrephes in European lakes, which could be due to higher abundance of available zooplankton prey or lower levels of planktivory in combination with the greater conspicuousness of Bythotrephes (Foster and Sprules, 2009). In Lake Michigan, the decline of D. retrocurva and D. pulicaria populations in contrast to relatively stable populations of D. mendotae has been attributed to the latter’s markedly faster escape responses (Pichlová-Ptácníková and Vanderploeg, 2011).

Some studies have documented an increase in chlorophyll a abundance with the invasion of Bythotrephes, potentially due to a release from grazing following increased predation pressure on zooplankton species (Barbiero and Rockwell, 2009; Hovius et al., 2007). This would imply that Bythotrephes is capable of affecting multiple trophic levels of the freshwater community. However, an increase in Bythotrephes abundance is not always correlated with an increase in chlorophyll a (Foster and Sprules, 2009; Strecker and Arnott, 2008). Notably, Strecker and Arnott (2008) demonstrated that invaded lakes experienced a significant reduction in secondary production, and hence a reduction in resources available in the epilimnion.

In some cases, Bythotrephes has been associated with a shift in cladoceran communities towards larger taxa over small possibly due to Bythotrephes predation of smaller species (Barbiero and Rockwell 2008; Hovius et al. 2007; Yan and Pawson 1997). Vertical migration has also been observed in Daphnia spp. and copepods (e.g., Diacyclops thomasi, Leptomdiaptomus ashlandi, L. minutus) populations following Bythotrephes invasion, indicating that some species may migrate to deeper waters during the day to avoid Bythotrephes predation or competition (Bourdeau et al. 2011; Jokela et al. 2011; Lehman and Cáceres 1993; Arwine et al. 2025). Diel vertical migration may lead to an indirect negative effect on these native prey species, including reduced individual and population growth rates at lower temperatures (Pangle and Peacor, 2006; Pangle et al. 2007).

Potential:
Significant increases in the trophic position of zooplankton (reduced herbivorous cladoceran biomass and increased omnivorous/predatory copepod biomass) and planktivorous fish such as the lake herring (Coregonus artedi) with the introduction of Bythotrephes has the potential to lead to substantial contaminant biomagnification in consumers (Rennie et al., 2011). However, increased mercury concentrations in consumers has not been observed following invasion; this may be attributed to increased foraging and growth efficiencies of consumers or to changes in the feeding habits of omnivorous prey (Rennie et al., 2011).
 

There is little or no evidence to support that Bythotrephes longimanus has significant socio-economic impacts in the Great Lakes.

The tail spines of Bythotrephes hook on fishing lines, fouling fishing gear. This problem has largely been eliminated with a switch to line/gear types less susceptible to Bythotrephes fouling.

There is little or no evidence to support that Bythotrephes longimanus has significant beneficial effects in the Great Lakes.

Bythotrephes is a food source for fish including yellow perch, white perch, walleye, white bass, alewife, bloater chub, Chinook salmon, emerald shiner, spottail shiner, rainbow smelt, lake herring, lake whitefish, and deepwater sculpin (Branstrator and Lehman 1996; Bur et al. 1986; Makarewicz and Jones 1990). However due to its long tail spine, predation of Bythotrephes is mainly restricted to larger fish and non-gape limited species (Pothoven et al. 2007).


Management: Regulations (pertaining to the Great Lakes)
In Wisconsin, the spiny waterflea is a prohibited invasive species (Wis. Admin. Code § NR 40.04), which indicates that it is likely to survive and spread if introduced into the state, potentially causing economic or environmental harm or harm to human health (Wis. Admin. Code § NR 40.02). With certain exceptions, it is unlawful to transport, possess, transfer or introduce a prohibited invasive species in Wisconsin (Wis. Admin. Code § NR 40.04). In Minnesota, the spiny waterflea is a regulated invasive species (Min. Admin. Rules § 6216.0260). It is legal to possess, sell, buy, and transport regulated invasive species, but no person may introduce a regulated invasive species without a permit (Min. Admin. Rules § 6216.0265 Subpart 1).

Note: Check federal, state/provincial, and local regulations for the most up-to-date information.

Control
Like the confamilial fishhook waterflea (Cercopagis pengoi), the spiny waterflea is most likely to be spread on aquatic equipment, especially fishing lines. Consequently, public education is a significant method of control which can greatly reduce incidences of species transfer by unaware or incautious anglers (Jacobs and MacIsaac, 2007; Lui et al., 2010).
 

Biological
Bythotrephes longimanus is consumed by rainbow smelt, lake herring, lake whitefish, yellow perch, white perch, white bass, pumpkinseed sunfish, walleye, alewife, bloater chub, emerald shiner, spottail shiner, deepwater sculpin, and chinook salmon in the Great Lakes (Bur et al. 1986; Makarewicz and Jones 1990; Branstrator and Lehman 1996; LeDuc et al. 2020 ). Bythotrephes longimanus’ defensive tailspine has been observed increasing in size throughout the summer in response to predation pressure (Straile and Halbich, 2000). Consequently, larger fish are more likely to be successful predators (Branstrator and Lehman 1996).  The opossum shrimp (Mysis relicta) has been observed eating B. longimanus in Ontario lakes, but the frequency of consumption appeared related to abundance of the invader and alternate prey (Nordin 2008).

Physical
Bythotrephes longimanus collects in gelatinous clumps on fishing lines, downrigger cables, and other aquatic equipment (Lui et al. 2010). Responsible maintenance and cleaning methods are recommended to prevent spread between water bodies, including cleaning all aquatic equipment with high pressure (>250 psi) or hot (>50°C) water after each use (Ontario’s Invading Species Awareness Program). The acute upper lethal temperature level for B. longimanus, at which death occurs rapidly, is 40°C (GLMRIS 2012), and a study found that B. longimanus specifically requires 10 minutes treatment with 43°C water to ensure 100% mortality (Beyer et al. 2011). Additional recommendations include air drying aquatic equipment for more than six hours to kill any potential spiny waterflea eggs (Minnesota DNR 2026).  Fishing lines designed specifically to prevent the spread of waterfleas, such as the Flea Flicker brand, have been proven effective in significantly reducing fouling on lines, indicating their importance as a management tool (Jacobs and MacIsaac 2007).


Electron beam irradiation has been used to control microorganisms in aquatic pathways, including Bythotrephes longimanus (GLMRIS 2012). Electron beam irradiation is a non-selective control method that exposes water to low doses of radiation using gamma-sterilizers or electron accelerators, breaking down DNA in living organisms while leaving behind no by-products (GLMRIS 2012). Ultraviolet (UV) light can also effectively control microorganisms, including B. longimanus, in water treatment facilities and narrow channels, where UV filters can be used to emit UV light into passing water, penetrating cell walls and rearranging DNA of microorganisms (GLMRIS 2012).

Chemical
There are no known chemical control methods for this species

Note: Check state/provincial and local regulations for the most up-to-date information regarding permits for control methods. Follow all label instructions.

This should include case studies for management, summarize management literature, etc.  Research on new tools for management is appropriate even if approvals for use are pending.  Do not use ‘trade names’ for any products (e.g., glyphosate is OK, Roundup is not).  Tone generally is toolbox rather than specific recommendations (which may depend on jurisdiction and specific situation).  Link directly to Collaboratives that work on management (assuming a link to a stable website is available) and/or provide management advice.


Remarks: Deweese et al (2021) report on preliminary microfossil evidence for the presence of Bythotrephes in 4 inland lakes of the Great Lakes region which would place the introduction dates for this species significantly earlier.  The authors still speculate whether the data may be an artefact (albeit of a common method), and should be considered preliminary at best. If these corroborated, earliest dates of reported microfossils should be 1934 (Lake Kabetogama, MN), 1908 (Mille Lacs, MN), 1900 (Lake Nipissing, Ontario) and 1650s (Three Mile Lake, Ontario).  Such early records call into question both the presumed means of introduction for this species (ballast water) as well as the possibility that the species should be considered native.


References (click for full reference list)


Author: Liebig, J., A. Benson, J. Larson, T.H. Makled, A. Fusaro, and K.M. Reaver


Contributing Agencies:
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Revision Date: 4/2/2026


Citation for this information:
Liebig, J., A. Benson, J. Larson, T.H. Makled, A. Fusaro, and K.M. Reaver, 2026, Bythotrephes longimanus: U.S. Geological Survey, Nonindigenous Aquatic Species Database, Gainesville, FL, and NOAA Great Lakes Aquatic Nonindigenous Species Information System, Ann Arbor, MI, https://nas.er.usgs.gov/queries/GreatLakes/FactSheet.aspx?Species_ID=162, Revision Date: 4/2/2026, Access Date: 4/14/2026

This information is preliminary or provisional and is subject to revision. It is being provided to meet the need for timely best science. The information has not received final approval by the U.S. Geological Survey (USGS) and is provided on the condition that neither the USGS nor the U.S. Government shall be held liable for any damages resulting from the authorized or unauthorized use of the information.