Percina shumardi (Girard, 1859)

Common Name: River Darter

Synonyms and Other Names:

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Identification: The body of the River Darter Percina shumardi is elongate and slightly compressed laterally. The back is brown to olive brown in color with lighter sides and a white belly. The snout is curved downwards and short. The mouth is slightly slanted and jaw protrudes past the eyes. River Darters are distinguished by 6-7 saddle marks on its back, 8-11 vertical bars along its side, and a black bar below the eyes. The spiny dorsal fin has a large black spot on tissue behind the last 3-4 spines. The caudal fins and second dorsal fins are lightly speckled and barred. Pelvic, pectoral, and anal fins are transparent to lightly pigmented. The species exhibits sexual dimorphism, males will normally have a line of modified scales from the anal fin to pelvic fin. Males will also have a thickened membrane on the edge of pelvic fins, while females lack this feature. Breeding males are typically darker and have more pronounced colors than females (Becker 1983).


Size: 7.8 cm.


Native Range: Hudson Bay basin, Ontario, Manitoba, North Dakota, and south in Great Lakes and Mississippi River basins to Louisiana; Gulf drainages from Mobile Bay, Alabama, to Neches River, Texas; isolated population in San Antonio drainage, Texas (Page and Burr 1991). Native to Lake Michigan, Lake Huron, Lake Erie. Absent from Lake Superior and Lake Ontario (Roth et al. 2013; Cudmore-Vokey and Crossman 2000). Although the species has been recorded from the Tennessee drainage in northeastern Tennessee, it has not been reported in Virginia (Jenkins and Burkhead 1994), North Carolina (Menhinick 1991), or Georgia (Dahlberg and Scott 1971a).


Great Lakes Nonindigenous Occurrences: The River Darter was collected from the city of Chicago's water system prior to 1926. The city's water source, and hence the source of the fish, was off-shore in Lake Michigan,  (Becker 1983).  Becker (1983) lists this species as introduced into the Lake Michigan drainage in the Wolf and Waupaca rivers and in lakes Winnebago and Poygan.  


Table 1. Great Lakes region nonindigenous occurrences, the earliest and latest observations in each state/province, and the tally and names of HUCs with observations†. Names and dates are hyperlinked to their relevant specimen records. The list of references for all nonindigenous occurrences of Percina shumardi are found here.

Full list of USGS occurrences

State/ProvinceFirst ObservedLast ObservedTotal HUCs with observations†HUCs with observations†
IL192619261Lake Michigan
WI196319832Lake Winnebago; Wolf

Table last updated 5/2/2024

† Populations may not be currently present.


Ecology: The River Darter inhabits deep riffles and chutes of medium to large rivers, in areas of moderate current and coarse gravel to rock substrates (Gilbert 1980). It is more frequently found in smaller streams during winter and spawning season in early spring. The species is also reportedly tolerant of stream impoundments (Etnier and Starnes 1993). River Darters can also be found in lakes along wave-swept shores with sand, gravel, or rubble, to depth of ~1 m (Becker 1983).

Spawning can occurs at 10°C and has been observed in February at the earliest. Larvae take 6-7 days to hatch at temperatures of 22°C and likely take longer to hatch at lower temperatures. Larvae are active during the day can be found drifting downstream near the surface. Once larvae have grown from the initial size of 4mm to 12-15mm they are considered juveniles. Juveniles are found in areas with sand or gravel substrate and moderate current (Etnier and Starnes 1993). Sexual maturity is reached at age 1 and the average lifespan of individuals is three years old (Dalton 1990).

River Darters are benthic invertivores, primarily consuming chironomids, trichopterans, and small crustaceans (Thomas 1970). Small zooplankton are the primary food source for juveniles (Becker 1983). It is also known to exploit pleurocerid snail populations (Haag and Warren 2006).


Great Lakes Means of Introduction: The River Darter may have found its way to Illinois waters of Lake Michigan through the Chicago Sanitary and Ship Canal. The species likely gained access to the Lake Michigan drainage in Wisconsin via the Wisconsin-Fox Canal at Portage, Wisconsin (Becker 1983). It was first reported from this area in 1963 from the western shore of Lake Winnebago (Becker 1983). This species was not found in this drainage in surveys conducted in the late 1920s (Becker 1983). See Remarks section for a possible alternate scenario.


Great Lakes Status: Cryptogenic. Currently considered native to Lake Huron (Bailey et al 2004) and western Lake Erie and non-native in Lake Michigan; but the native range for the region is discontinuous with other populations. Overwintering and reproducing, but unquestionably rare (Mandrak 2017, personal communication). COSEWIC (2016) designated populations of the species in the Great Lakes – Upper St. Lawrence as endangered.


Great Lakes Impacts: Current research on the environmental impact of Percina shumardi in the Great Lakes is inadequate to support proper assessment.

The River Darter is largely limited to large rivers (Thomas 1970) and is therefore unlikely to have impacts in the open lake, though it may impact biotic communities in the connecting channels and river mouths.   Hybridization is reportedly rare for the genus (Thomas 1970).

There is little or no evidence to support that the Percina shumardi has significant socio-economic impacts in the Great Lakes.

There is little or no evidence to support that Percina shumardi has significant beneficial impacts in the Great Lakes.

Percina shumardi is of little use as a bait fish or forage fish (Becker 1983)


Management: Regulations

There are no known regulations for this species.

Note: Check federal, state, and local regulations for the most up-to-date information

Control
Biological

There are no known biological control methods for this species.

Physical

Success of physical control methods have not been studied on the River Darter.  General methods of fish control include accelerated water velocity, physical barriers, pressurized hot water/steam, hot water thermal barriers , removal of catch limits, reservoir drawdowns, traps, nets, electrofishing, and combinations of treatments (GLMRIS 2011, Meronek et al. 1996). Patrick et al. (1985) observed that air bubble curtains have been successful in deterring various species of fish—especially when used in conjunction with strobe lights.

Chemical

Chemical piscicides antimycin A and rotenone are general piscicides, the use of which has not been studied on The River Darter. These piscicides are toxic to other species and can cause non-target kills (GLMRIS 2011). Careful consideration should be taken with chemical methods to reduce impact to non-target species.

Note: Check state/provincial and local regulations for the most up-to-date information regarding permits for control methods. Follow all label instructions.


Remarks: Becker (1983) described the low divide between the Fox and Wisconsin rivers at Portage as one of the major historical natural crossover connections between the Mississippi River and Great Lakes drainage basins. Historical reports from both before and after construction of the Portage Canal describe movement of water from the Wisconsin River through the divide to the Fox River; hydrologic analysis combined with local topography indicate that this connection has been a regular historical occurrence (J. Lyons, Wisconsin DNR, personal communication). It is likely that P. shumardi is native to the Fox River drainage, having colonized from the Wisconsin River at some point since the last glaciation, and was simply overlooked prior to Becker (1983) due to the higher sampling intensity of Becker et al. in the Fox River relative to earlier collectors and the low abundance of P. shumardi in the Fox River drainage (J. Lyons, Wisconsin DNR, personal communication).

This species should be considered cryptogenic in Lakes Huron, Michigan and Erie. Percina shumardi is  considered native to Lake Huron according to Bailey et al. 2004 (Atlas of Michigan Fishes), and questionably native by Cudmore-Vokey and Crossman (2000).  Roth et al (2011) considered it native to these three lakes, likely endangered and declining; but Roth independently wrote that it "may be invasive to Lake Michigan, although it is listed within the range proposed by Scott and Crossman and Hubbs et al. 2004.  It was considered possibly introduced to Lake Michigan through a canal (Bailey & Smith 1981, Becker 1983, but see Hubbs & Lagler, Roth et al. in prep.). The species is unquestionably rare in the Great Lakes” [Mandrak 2017, personal communication].


References (click for full reference list)


Author: Pam Fuller, Matt Neilson and Kylan Hopper


Contributing Agencies:
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Revision Date: 1/17/2024


Peer Review Date: 6/5/2012


Citation for this information:
Pam Fuller, Matt Neilson and Kylan Hopper, 2024, Percina shumardi (Girard, 1859): U.S. Geological Survey, Nonindigenous Aquatic Species Database, Gainesville, FL, and NOAA Great Lakes Aquatic Nonindigenous Species Information System, Ann Arbor, MI, https://nas.er.usgs.gov/queries/greatLakes/FactSheet.aspx?Species_ID=826&Potential=N&Type=0&HUCNumber=DGreatLakes, Revision Date: 1/17/2024, Peer Review Date: 6/5/2012, Access Date: 5/2/2024

This information is preliminary or provisional and is subject to revision. It is being provided to meet the need for timely best science. The information has not received final approval by the U.S. Geological Survey (USGS) and is provided on the condition that neither the USGS nor the U.S. Government shall be held liable for any damages resulting from the authorized or unauthorized use of the information.