Reference List

This list includes references used to derive specimen records as well as those with scientific name Hemimysis anomala listed in key words.

Results also available in (click to export table to spreadsheet)

REFTypeCut-and-Paste Reference
32155 Report Anonymous. 2018. Lake Superior Research Institute documents non-native zooplankton in Western Lake Superior. University of Wisconsin Superior, Superior, WI.
13589 Journal Article Bij de Vaate, A., K. Jażdżewski, H.A.M. Ketelaars, S. Gollasch, and G. van der Velde. 2002. Geographical patterns in range expansion of Ponto-Caspian macroinvertebrate species in Europe. Canadian Journal of Fisheries and Aquatic Sciences 59(7):1159–1174. https://doi.org/10.1139/f02-098.
15936 News Black, D. 2007. Big trouble from little shrimp? Toronto Star. 2007 (January 20). http://www.thestar.com/News/article/173074 (accessed January 22, 2007).
34233 Journal Article Bodis, E., P. Borza, I. Potyo, M. Puky, A. Weiperth, and G. Guti. 2012. Invasive mollusc, crustacean, fish and reptile species along the Hungarian stretch of the River Danube and some connected waters. Acta Zoologica Academiae Scientiarum Hungaricae 58(Suppl):29–45.
44880 Report Boivin, V., N. Vachon, and P. Brodeur. 2022. Régime alimentaire des poissons capturés dans le cadre du Réseau de suivi ichtyologique du fleuve Saint-Laurent. Secteurs lac Saint-Pierre et Bécancour-Batiscan 2021. Ministère de l'Environnement, de la Lutte contre les changements climatiques, de la Faune et des Parcs, Quebec, Canada.
44881 Report Boivin, V., P. Brodeur, and N. Vachon. 2024. Régime alimentaire des poissons capturés dans le cadre du Réseau de suivi ichtyologique du fleuve Saint-Laurent. Secteur Montréal-Sorel en 2023. Ministère de l'Environnement, de la Lutte contre les changements climatiques, de la Faune et des Parcs, Quebec, Canada.
16400 Book Chapter Bondarenko, M.V., and E.A. Yablonskaya. 1979. The rate of oxygen consumption by the mysid Paramysis lacustris (Czern.) from the northern Caspian Sea. Page 65–69 in Effect of water management on the biological productivity of the Azov and Caspian seas. VNIRO. Moscow, Russia.
16401 Journal Article Borcherding, J., S. Murawski, and H. Arndt. 2006. Population ecology, vertical migration and feeding of the Ponto-Caspian invader Hemimysis anomala in a gravel-pit lake connected to the River Rhine. Freshwater Biology(51):2376–2387.
36817 Journal Article Borza, P. 2014. Life history of invasive Ponto-Caspian mysids (Crustacea: Mysida): A comparative study. Limnologica 44. https://doi.org/10.1016/j.limno.2013.06.001.
46675 Journal Article Branstrator, D.K., K. Huelsbeck, G.M. Thelen, B. Johnson, M.A. Nagel, L.R. Isbell, L.M. Fanberg, H. Schaefer, M.E. Brown, and M.C. TenEyck. 2026. First evidence of Hemimysis anomala (bloody red shrimp) establishment in Lake Superior. Journal of Great Lakes Research 52(2):102779. https://doi.org/10.1016/j.jglr.2026.102779.
24646 Journal Article Brooking, T.E., L.G. Rudstam, S.D. Krueger, J.R. Jackson, A.B. Welsh, and W.W. Fetzer. 2010. First occurrence of the mysid Hemimysis anomala in an inland lake in North America, Oneida Lake. Journal of Great Lakes Research 36(3):577–381.
39198 Journal Article Brown, M.E., B.T. Boscarino, J.Q.G. Roellke, E.T.K. Stapylton, and A.R. Driller-Colangelo. 2014. Fifteen miles on the Erie Canal: the spread of Hemimysis anomala G.O. Sars, 1907 (bloody red shrimp) in the New York State canal system. International Journal of Field Research on Biological Invasions 3(4):261–267. https://doi.org/10.3391/bir.2014.3.4.07.
46087 Journal Article Brown, M.E., K.L. Buffington, L.B. Cleckner, and N.R. Razavi. 2022. Elevated methylmercury concentration and trophic position of the non-native bloody red shrimp (Hemimysis anomala) increase biomagnification risk in nearshore food webs. Journal of Great Lakes Research 48(1):252–259. https://doi.org/10.1016/j.jglr.2021.11.010.
16402 Journal Article Bubinas, A.D. 1980. Formation of benthic fauna as a food base for fish in the reservoir of the Kaunas hydro electric power plant, Lithuanian-SSR USSR. Lietuvos TSR Mokslu Akademijos Darbai Serija C Biologijos Mokslai(4):91–96.
16403 Journal Article Cabana, G., A. Tremblay, J. Kalff, and J.B. Rasmussen. 1994. Pelagic food chain structure in Ontario lakes: a determinant of mercury levels in lake trout (Salvelinus namaycush). Canadian Journal of Fisheries and Aquatic Sciences 51:381–389.
46224 Journal Article Cangelosi, A., M. Balcer, K. Prihoda, M. Gruwell, M. TenEyck, R. Aicher, Y. Lopez-Camacho, I.T. Knight, and E.K. Grey. 2024. Evaluation of eDNA qPCR monitoring as an early detection tool for a non-native mysid in Great Lakes Waters. Journal of Great Lakes Research 50(4):102377. https://doi.org/10.1016/j.jglr.2024.102377.
45035 Journal Article Coughlan, N.E., S. O'Hara, K. Crane, J.T.A. Dick, H.J. MacIsaac, and R.N. Cuthbert. 2020. Touch too much: aquatic disinfectant and steam exposure treatments can inhibit further spread of invasive bloody-red mysid shrimp Hemimysis anomala. Wetlands Ecology and Management 28:397–402. https://link.springer.com/article/10.1007/s11273-020-09720-5.
44907 Journal Article Dickey, J.W.E., J.M. Jeschke, M. Lentz, E. Briski, E. Kazanaviciute, S.A. Berger, and J.C. Nejstgaard. 2024. First occurrence of the mysid Hemimysis anomala G.O. Sars, 1907 in Lake Stechlin, Germany. BioInvasions Records 13(4):927–940. https://www.reabic.net/journals/bir/2024/4/BIR_2024_Dickey_etal.pdf.
16404 Journal Article Dumont, S. 2006. A new invasive species in the north-east of France, Hemimysis anomala G. O. Sars, 1907 (Mysidacea). Crustaceana 79(10):1269–1274. https://doi.org/10.1163/156854006778859560.
24647 Journal Article Dumont, S., and C.D. Muller. 2009. Distribution, ecology and impact of a small invasive shellfish, Hemimysis anomala in Alsatian water. Biological Invasions 12(3):495–500.
26306 Journal Article Ellis, S., and H.J. MacIsaac. 2009. Salinity tolerance of Great Lakes invaders. Freshwater Biology 54(1):77–89.
32123 Journal Article Evans, T.M., R. Naddafi, B.C. Weidel, B.F. Lantry, M.G. Walsh, B.T. Boscarino, O.E. Johannsson, and L.G. Rudstam. 2018. Stomach contents and stable isotopes analysis indicate Hemimysis anomala in Lake Ontario are broadly omnivorous>. Journal of Great Lakes Research 44(3):467–475. https://www.sciencedirect.com/science/article/pii/S0380133018300388.
34237 Journal Article Gallardo, B., and D.C. Aldridge. 2015. Is Great Britain heading for a Ponto–Caspian invasional meltdown? Journal of Applied Ecology 52:41–49. https://doi.org/10.1111/1365-2664.12348.
26239 Report GLMRIS. 2012. Appendix C: Inventory of available controls for aquatic nuisance species of concern, Chicago Area Waterway System. U.S. Army Corps of Engineers. http://glmris.anl.gov/documents/docs/anscontrol/All27ANSControlFactSheets.pdf.
30785 Database Harris, B. 2017. Collection data on non-native aquatic species in Lake Michigan. U.S. Fish and Wildlife Service - Green Bay Fish and Wildlife Conservation Office, Green Bay, WI. Accessed on 11/16/2017.
15922 Journal Article Holdich, D., S. Gallagher, L. Rippon, P. Harding, and R. Stubbington. 2006. The invasive Ponto-Caspian mysid, Hemimysis anomala, reaches the UK. Aquatic Invasions 1(1):4–6.
27712 Database iMapInvasives: NatureServe’s online data system supporting strategic invasive species management. 2019. New York iMapInvasives. © NatureServe. https://www.imapinvasives.org. Accessed on 07/08/2015.
16406 Journal Article Janas, U., and P. Wysocki. 2005. Hemimysis anomala G. O. Sars, 1907 (Crustacea, Mysidacea) – first record in the Gulf of Gdansk. Oceanologia 47(3):405–408.
16407 Journal Article Jażdżewski, K., A. Konopacka, and M. Grabowski. 2005. Native and alien malacostracan crustacean along the Polish Baltic Sea coast in the twentieth century. Oceanological and Hydrobiological Studies 24. https://www.researchgate.net/publication/215701058_Native_and_alien_Malacostracan_Crustacea_along_the_Polish_Baltic_Sea_coast_in_the_twentieth_century.
18337 Journal Article Kestrup, Å.M., and A. Ricciardi. 2008. Occurrence of the Ponto-Caspian mysid shrimp Hemimysis anomala (Crustacea, Mysida) in the St. Lawrence River. Aquatic Invasions 3(4):461–464. http://www.aquaticinvasions.net/2008/AI_2008_3_4_Kestrup_Ricciardi.pdf.
24648 Journal Article Kestrup, Å.M., J.T.A. Dick, and A. Ricciardi. 2011. Interactions between invasive and native crustaceans: differential functional responses of intraguild predators towards juvenile hetero-specifics. Biological Invasions 13(3):731–737.
16408 Journal Article Ketelaars, H.A.M., F.E. Lambreqts-van de Clundert, C.J. Carpentier, A.J. Waqenvoort, and W. Hooqenboezem. 1999. Ecological effects of the mass occurrence of the Ponto-Caspian invader, Hemimysis anomala G.O. Sars, 1907 (Crustacea: Mysidacea), in a freshwater storage reservoir in the Netherlands, with notes on its autecology and new records. Hydrobiologia 394:233–248. https://doi.org/10.1023/A:1003619631920.
16409 Journal Article Komarova, T.I. 1989. Ecological and faunistic review of Mysidacea (Crustacea: Mysidae) in the Sea of Azov, USSR. Vestnik Zoologii(4):3–7.
16410 Journal Article Langeland, A., J.I. Koksvik, and J. Nydal. 1991. Impact of the introduction of Mysis relicta on the zooplankton and fish populations in a Norwegian lake. American Fisheries Society Symposium(9):98–114.
19528 Journal Article Lantry, B.F., M.G. Walsh, J.H. Johnson, and J.E. McKenna Jr. 2010. Occurrence of the Great Lake's most recent invader, Hemimysis anomala, in the diet of fishes in southeastern Lake Ontario. Journal of Great Lakes Research 36:179–183.
16411 Journal Article Lasenby, D.C., T.G. Northcote, and M. Fürst. 1986. Theory, practice, and effects of Mysis relicta introductions to North American and Scandinavian lakes. Canadian Journal of Fisheries and Aquatic Sciences 43:1277–1284.
16412 Journal Article Lundberg, S., and J.-E. Svensson. 2004. The mysid shrimp Hemimysis anomala G. O. Sars documented in Sweden. Fauna och Flora (Stockholm) 99(1):38–40.
26304 Journal Article Marty, J., K. Bowen, M.A. Koops, and M. Power. 2010. Distribution and ecology of Hemimysis anomala, the latest invader of the Great Lakes basin. Hydrobiologia 647(1):71–80.
46130 Database Ministère de l'Environnement, de la Lutte contre les changements climatiques, de la Faune et des Parcs (MELCCFP). 2025. Invasive alien aquatic fauna in Quebec, updated 2025-06-10. Government of Québec, Montreal, Quebec, Canada. https://www.donneesquebec.ca/recherche/dataset/faune-aquatique-exotique-envahissante-au-quebec]. Created on 01/11/2024. Accessed on 07/17/2025.
16413 Journal Article Mordukhai-Boltovskoi, P.D. 1979. Composition and distribution of Caspian fauna in light of modern data. Internationale Revue der gesamten Hydrobiologie 64:1–38. https://doi.org/10.1002/iroh.19790640102.
45006 Database NatureServe. 2025. iMapInvasives: NatureServe’s online data system supporting strategic invasive species management. NatureServe. http://www.imapinvasives.org. Accessed on 03/18/2025.
16414 Journal Article Northcote, T.G. 1991. Success, problems, and control of introduced mysid populations in lakes and reservoirs. American Fisheries Society Symposium(9):5–16.
16415 Journal Article Olenin, S., and E. Leppäkoski. 1999. Non-native animals in the Baltic Sea: alteration of benthic habitats in coastal inlets and lagoons. Hydrobiologia 393:233–243. https://link.springer.com/chapter/10.1007%2F978-94-017-0912-5_24.
19331 Database Ontario Federation of Anglers and Hunters. 2009. OFAH Database All Species - 2009 Download.
33370 Web Page Ontario's Invading Species Awareness Program. Bloody Red Shrimp Hemimysis anomala. http://www.invadingspecies.com/invaders/invertebrates/bloody-red-shrimp/. Accessed on 06/27/2013.
33829 Web Page Pennsylvania State University Behrend. 2019. Behrend students find new species in Lake Erie. https://behrend.psu.edu/feature/behrend-students-find-new-species-lake-erie. Created on 01/01/2019. Accessed on 10/29/2019.
26891 Journal Article Perez-Fuentetaja, A., and J. Wuerstle. 2014. Prey size selection and feeding ecology of an omnivorous invader: Hemimysis anomala. Journal of Great Lakes Research 40(2):257–264.
16416 Journal Article Pienimäki, M.E., and E. Leppäkoski. 2004. Invasion pressure on the Finnish Lake District: invasion corridors and barriers. Biological Invasions(6):331–346.
16417 Journal Article Pligin, Y.V., and L.V. Yemel'yanova. 1989. Acclimatization of Caspian invertebrates in Dnieper reservoirs. Hydrobiological Journal(25):1–9.
16418 Journal Article Pothoven, S.A., I.A. Grigorovich, G.L. Fahnenstiel, and M.D. Balcer. 2007. Introduction of the Ponto-Caspian bloody-red mysid Hemimysis anomala into the Lake Michigan basin. Journal of Great Lakes Research 33:285–292. https://doi.org/10.3394/0380-1330(2007)33[285:IOTPBM]2.0.CO;2.
16419 Journal Article Rasmussen, J.B., Rowan, D.J., Lean, D.R.S., and Carey, J.H. 1990. Food chain structure in Ontario lakes determines PCB levels in lake trout (Salvelinus namaycush) and other pelagic fish. Canadian Journal of Fisheries and Aquatic Sciences 47:2030–2038.
16420 Journal Article Ricciardi, A., and J.B. Rasmussen. 1998. Predicting the identity and impact of future biological invaders: a priority for aquatic resource management. Canadian Journal of Fisheries and Aquatic Sciences 55(7):1759–1765.
24649 Journal Article Ricciardi, A., S. Avlijas, and J. Marty. 2011. Forcasting the ecological impacts of the Hemimysis anomala invasion in North America: Lessons from other freshwater mysid introductions. Journal of Great Lakes Research 38(2):7–13.
46189 Journal Article Rogissart, H., V. Frossard, J. Guillard, C. Rautureau, and S. Jacquet. 2024. Tracking the real-time behavior of Hemimysis anomala’s winter swarms using acoustic camera. Journal of Great Lakes Research 50(1):102250. https://doi.org/10.1016/j.jglr.2023.102250.
16421 Journal Article Salemaa, H., and V. Hietalahti. 1993. Hemimysis anomala G. O. Sars (Crustacea: Mysidacea) – immigration of a Pontocaspian mysid into the Baltic Sea. Annales Zoologici Fennici 30(4):271–276.
44978 Journal Article Sheehan, R., I.T. Knight, A. Cangelosi, A. Melendez, H. Phillips, A. Welsbacher, and M.E. Gruwell. 2024. Quantitative assessment of planktonic AIS eDNA signal duration in Great Lakes harbor water microcosm. Management of Biological Invasions 15(4):567–580. https://www.reabic.net/journals/mbi/2024/4/MBI_2024_ICMB_Sheehan_etal.pdf.
30968 Journal Article Sinclair, J. S., Arnott, S. E., and A. Cox. The quick and the dead: copepods dominate as cladocerans decline following invasion by Hemimysis anomala. Canadian Journal of Fisheries and Aquatic Sciences 73:793–803.
30945 News Smith, P. A. 2017. Invasive bloody red shrimp's impact on ecosystem unknown. Milwaukee Journal Sentinel. Milwaukee, WI. https://www.jsonline.com/story/sports/outdoors/2017/08/02/smith-invasive-bloody-red-shrimps-impact-ecosystem-unknown/502884001/. Created on 08/02/2017. Accessed on 01/25/2018.
16422 Journal Article Spencer, C.N., B.R. McClelland, and J.A. Stanford. 1991. Shrimp stocking, salmon collapse, and eagle displacement. BioScience 44:14–21.
20785 Journal Article Sprung, M. 1995. Physiological energetics of the zebra mussel Dreissena polymorpha in lakes. I: Growth and reproductive effort. Hydrobiologia 304(2):117–132.
39842 Database U.S. Fish and Wildlife Service. 2020. Benthic Macroinvertebrate AIS Early Detection efforts on Lake Superior. U.S. Fish and Wildlife Service, Ashland, WI. Accessed on 09/16/2022.
46239 Journal Article Wachala, S.N., J. Janssen, E.B. Young, and J.A. Berges. 2025. Seasonal population characteristics and gut contents of the invasive mysid, Hemimysis anomala, in Milwaukee Harbor, Lake Michigan. Journal of Great Lakes Research 51(1):102462. https://doi.org/10.1016/j.jglr.2024.102462.
19502 Journal Article Walsh, M.G., B.F. Lantry, B. Boscarino, K. Bowen, J. Gerlofsma, T. Schaner, R. Back, J. Questel, A.G. Smythe, R. Cap, M. Goehle, B. Young, M. Chalupnicki, J.H. Johnson, and J.E. McKenna. 2010. Early observations on an emerging Great Lakes invader Hemimysis anomala in Lake Ontario. Journal of Great Lakes Research 36(3):499–504.
37839 Journal Article Walsh, M.G., B.T. Boscarino, J. Marty, and O.E. Johannsson. 2012. Mysis diluviana and Hemimysis anomala: Reviewing the roles of a native and invasive mysid in the Laurentian Great Lakes region. Journal of Great Lakes Research 38. https://doi.org/10.1016/j.jglr.2012.03.003.
16423 Conference Proceedings Wittman, K.J. 2006. Distribution and invasion potential of the Ponto-Capsian Mysidae (Crustacea: Malacostraca: Peracarida: Mysida). Abstract only. Page 41 in Rabitsch, W., F. Klingenstein, and F. Essl, eds. Neobiota: from ecology to conservation. 4th European conference on biological invasions, 27-29 September 2006, Vienna, Austria. Bundesamt für Naturschutz. Bonn, Germany.
16424 Journal Article Wittman, K.J. 2007. Continued massive invasion of Mysidae in the Rhine and Danube river systems, with first records of the order Mysidacea (Crustacea: Malacostraca: Peracarida) for Switzerland. Revue Suisse de Zoologie 114(1):1–22.
36833 Journal Article Wittmann, K.J. 2007. Continued massive invasion of Mysidae in the Rhine and Danube river systems, with first records of the order Mysidacea (Crustacea: Malacostraca: Peracarida) for Switzerland. Revue Suisse de Zoologie 114(1):1. https://doi.org/10.5962/bhl.part.80389.
36520 Journal Article Wittmann, K.J., and A.P. Ariani. 2009. Reappraisal and range extension of non-indigenous Mysidae (Crustacea, Mysida) in continental and coastal waters of eastern France. Biological Invasions 11(2):2. https://doi.org/10.1007/s10530-008-9257-7.
26830 Database Wunderlich, M. 2012. 2012 Eastern Finger Lakes Benthic Aquatic Invertebrate Assessment (unpublished). Created on 05/12/2014.
24659 Journal Article Yuille, M.J., T.B. Jonhson, S.E. Arnott, and L.M. Campbell. 2012. Hemimysis anomala in Lake Ontario food webs: Stable isotope analysis of nearshore communities. Journal of Great Lakes Research 38(Supplement 2):86–92.


Disclaimer:

The data represented on this site vary in accuracy, scale, completeness, extent of coverage and origin. It is the user's responsibility to use these data consistent with their intended purpose and within stated limitations. We highly recommend reviewing metadata files prior to interpreting these data.

Citation information: U.S. Geological Survey. [2026]. Nonindigenous Aquatic Species Database. Gainesville, Florida. Accessed [5/4/2026].

Contact us if you are using data from this site for a publication to make sure the data are being used appropriately and for potential co-authorship if warranted.

For general information and questions about the database, contact Wesley Daniel. For problems and technical issues, contact Matthew Neilson.