The Nonindigenous Occurrences section of the NAS species profiles has a new structure. The section is now dynamically updated from the NAS database to ensure that it contains the most current and accurate information. Occurrences are summarized in Table 1, alphabetically by state, with years of earliest and most recent observations, and the tally and names of drainages where the species was observed. The table contains hyperlinks to collections tables of specimens based on the states, years, and drainages selected. References to specimens that were not obtained through sighting reports and personal communications are found through the hyperlink in the Table 1 caption or through the individual specimens linked in the collections tables.

Pila scutata
(an applesnail)

Copyright Info
Pila scutata (Mousson, 1848)

Common name: an applesnail

Synonyms and Other Names: Pila conica

Native Range: Indonesia, Malaysia, Myanmar, Philippines, Cambodia, Laos, Vietnam, and Singapore (Low et al. 2013).

Hydrologic Unit Codes (HUCs) Explained
Interactive maps: Point Distribution Maps

Nonindigenous Occurrences:

Table 1. States with nonindigenous occurrences, the earliest and latest observations in each state, and the tally and names of HUCs with observations†. Names and dates are hyperlinked to their relevant specimen records. The list of references for all nonindigenous occurrences of Pila scutata are found here.

StateFirst ObservedLast ObservedTotal HUCs with observations†HUCs with observations†
HI196920064Hawaii Region; Maui; Molokai; Oahu

Table last updated 7/14/2024

† Populations may not be currently present.

Status: Established on Hawaii (Cowie et al. 2017) and Guam (Smith 1992, 2003).

References: (click for full references)

Cowie, R.H., K.A. Hayes, E.E. Strong, and S.C. Thiengo. 2017. Non-native apple snails: systematics, distribution, invasion history and reasons for introduction. Pages 3-32 in Joshi, R.C., Cowie, R.H., and Sebastian, L.S, eds. Biology and management of invasive apple snails. Philippine Rice Research Institute (PhilRice), Maligaya. Science City of Muñoz, Nueva Ecija 3119, 406 pp.

Low, M.E.Y., S.K. Tan, and T.H. Ng. 2013. Pila conica (Wood, 1828), or Pila scutata (Mousson, 1848)? The correct name for the native apple snail of Singapore (Gastropoda: Ampullariidae). Nature in Singapore 6:55-60.

Smith, B. D. 1992. Introduction and dispersal of apple snails (Ampullariidae) on Guam. Pacific Science Association Information Bulletin 44(1/2): 12–14.

Smith, B. D., 2003. Prosobranch gastropods of Guam. Micronesica 35–36: 244–270.

Author: Benson, A.J.

Revision Date: 7/3/2019

Citation Information:
Benson, A.J., 2024, Pila scutata (Mousson, 1848): U.S. Geological Survey, Nonindigenous Aquatic Species Database, Gainesville, FL, https://nas.er.usgs.gov/queries/FactSheet.aspx?SpeciesID=2258, Revision Date: 7/3/2019, Access Date: 7/14/2024

This information is preliminary or provisional and is subject to revision. It is being provided to meet the need for timely best science. The information has not received final approval by the U.S. Geological Survey (USGS) and is provided on the condition that neither the USGS nor the U.S. Government shall be held liable for any damages resulting from the authorized or unauthorized use of the information.


The data represented on this site vary in accuracy, scale, completeness, extent of coverage and origin. It is the user's responsibility to use these data consistent with their intended purpose and within stated limitations. We highly recommend reviewing metadata files prior to interpreting these data.

Citation information: U.S. Geological Survey. [2024]. Nonindigenous Aquatic Species Database. Gainesville, Florida. Accessed [7/14/2024].

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