Reference List

This list includes references used to derive specimen records.

Results also available in CSV (click to export table to spreadsheet)
Ref IDTypeAuthorDateTitle
46016 DatabaseAponte, N.0University of Puerto Rico Herbarium (MSM)
46118 ReportBecking, L.E., T. Bussel, M.S. Engel, M. Christianen, A.O. Debrot2014Proximate response of fish, conch, and sea turtles to the presence of the invasive seagrass Halophila stipulacea in Bonaire
46108 ReportBervoets, T.2011Results of Nature Foundation Research into invasive Seagrass H. stipulacea in the Simpson Bay Lagoon
46133 ReportBrubaker, M.2023Baseline assessment of invasive Halophila stipulacea in Virgin Islands National Park, St. John, USVI
45524 Journal ArticleCampbell, J.E., A. Allen, D.C. Sattelberger, M.D. White, and J.W. Fourqurean2025First record of the seagrass Halophila stipulacea (Forsskal) Ascherson in the waters of the continental United States (Key Biscayne, Florida)
45522 Journal ArticleCassell, J.S., E. Cruz-Rivera, S. Wyllie-Echeverria, and P. Jobsis2024Variation in nutritional quality of an invasive seagrass does not explain its low palatability to two key herbivores in a Caribbean Bay
46110 ReportDebrot, A.O., A. Hylkema, W. Vogelaar, H.W.G. Meesters, M.S. Engel, R. Leon, W.F. Prud'homme van Reine, and I. Nagelkerken2012Baseline surveys of Lac Bay benthic and fish communities, Bonaire
46433 Journal ArticleEbersole, A., J.E. Kaldy, C. Sullivan, A. Dieppa, I.C. Cosme, and M.O. Lugo2025Plant characteristics of the non-native seagrass Halophila stipulacea in recently colonized beds of Puerto Rico
46131 Journal ArticleFerry, R., Y. Buske, J. Poupin, and J. Smith-Ravin2017First record of the invasive swimming crab Charybdis hellerii (A. Milne Edwards, 1867) (Crustacea, Portunidae) off Martinique, French Lesser Antilles
46306 DatabaseFlorida Department of Environmental Protection2025Southeast Florida Action Network (SEAFAN)
46025 Journal ArticleHernández-Delgado, E.A., C. Toledo-Hernández, C.P. Ruíz-Díaz, N. Gómez-Andújar, J.L. Medina-Muñiz, M.F. Canals-Silander, and S.E. Suleimán-Ramos 2020Hurricane Impacts and the Resilience of the Invasive Sea Vine, Halophila stipulacea: a Case Study from Puerto Rico
25944 DatabaseiNaturalist2024
46434 Journal ArticleKaldy, J.E., C. Sullivan, A. Dieppa, E. Huertas, M. Reiss, I. Wojtenko, J. Perzley, I.C. Cosme, and M.O. Lugo2025Preliminary assessment of a nutrient pollution indicator for application to tropical seagrasses of Puerto Rico
46134 Journal ArticleKaldy, J.E., C. Sullivan, A. Dieppa, I. Cappielo Cosme, M. Orizondo Lugo, and M. Schiebout2023First record of phytomyxid infection of the non-native seagrass Halophila stipulacea in Puerto Rico
45510 Journal ArticleKolatkova, V., F.O.H. Smulders, E.A. Ward, and M. Vohnik2022Range expansion of Marinomyxa marina, a phytomyxid parasite of the invasive seagrass Halophila stipulacea, to the Caribbean
46119 Journal ArticleMaréchal, J.-P., E.H. Meesters, F. Vedie, and C. Hellio2013Occurrence of the alien seagrass Halophila stipulacea in Martinique (French West Indies)
45834 Journal ArticleOlinger L.K., S.L. Heidmann, A.N. Durdall, C. Howe, T. Ramseyer, S.G. Thomas, D.N. Lasseigne, E.J. Brown, J.S. Cassell, M.M. Donihe, M.D. Duffing Romero, M.A. Duke, D. Green, P. Hillbrand, K.R. Wilson Grimes, R.S. Nemeth, T.B. Smith, and M. Brandt2017Altered juvenile fish communities associated with invasive Halophila stipulacea seagrass habitats in the U.S. Virgin Islands
46140 Journal ArticleRodríguez-Jerez, Y., R. Sellares-Blasco, M.F. Villalpando, E. Vargas-Pérez, A. Valdez-Trinidad, S. Zambrano, Y.D. Evangelista, S.D. Guendulain-García, and A. Croquer2025First report of Halophila stipulacea (Forsskål) Ascherson,1867 in the Dominican Republic, an invasive species that continues to expand across the Caribbean
46015 Journal ArticleRuiz, H., and D.L. Ballantine2004Occurrence of the seagrass Halophila stipulacea in the tropical West Atlantic
45839 Journal ArticleRuiz, H., D.L. Ballantine, and J. Sabater2017Continued Spread of the seagrass Halophila stipulacea in the Caribbean: Documentation in Puerto Rico and the British Virgin Islands
46023 Journal ArticleSánchez-González, J.L., C.P. Ruiz-Díaz, C. Toledo-Hernández, and A.M. Sabat2025Effects of a Category One Hurricane on Caribbean Native and Invasive Seagrasses
45836 Journal ArticleScheibling, R.E., D.G. Patriquin, and K. Filbee-Dexter2018Distribution and abundance of the invasive seagrass Halophila stipulacea and associated benthic macrofauna in Carriacou, Grenadines, Eastern Caribbean
16075 DatabaseSmithsonian Institution2019United States National Herbarium (US)
45843 Journal ArticleSmulders, F.O.H., J.A. Vonk, M.S. Engel, and M.J.A. Christianen2017Expansion and fragment settlement of the non-native seagrass Halophila stipulacea in a Caribbean bay
46122 ReportSteiner, S.C.C., and D.A. Willette2013The invasive seagrass Halophila stipulacea (Hydrocharitaceae, Angiospermae) and its impact on the benthic landscape of Dominica, Lesser Antilles
46123 Journal ArticleSteiner, S.C.C., K.J. Macfarlane, L.M. Price, and D.A. Willette2010The distribution of seagrasses in Dominica, Lesser Antilles
46137 Journal ArticleValdez, S.R., E.C. Shaver, D.A. Keller, J.P. Morton, Y.S. Zhang, C. Wiernicki, C. Chen, C. Martinez, and B.R. Silliman2021A survey of benthic invertebrate communities in native and non-native seagrass beds in St. John, USVI
46127 Journal Articlevan Tussenbroek, B.I., M.M. van Katwijk, T.J. Bouma, T. van der Heide, L.L. Govers, and R.S.E.W. Leuven2016Non-native seagrass Halophila stipulacea forms dense mats under eutrophic conditions in the Caribbean
45521 Journal ArticleVaughn, K.M., A. Durdall, D.A. Willette, M. Brandt, S. Costa, and K.W. Grimes2024Sediment carbon storage in subtidal beds of the invasive seagrass Halophila stipulacea along an extreme water depth gradient, St. Thomas, U.S. Virgin Islands
46129 Journal ArticleVera, B., L. Collado-Vides, C. Moreno, and B.I. van Tussenbroek2014Halophila stipulacea (Hydrocharitaceae): A Recent Introduction to the Continental Waters of Venezuela
45853 Journal ArticleWaycott M., K.-j. van Dijk, A. Calladine, E. Bricker, and E. Biffin2021Genomics-Based Phylogenetic and Population Genetic Analysis of Global Samples Confirms Halophila johnsonii Eiseman as Halophila ovalis (R.Br.) Hook.f.
45835 Journal ArticleWillette, D.A., and R.F. Ambrose2009The distribution and expansion of the invasive seagrass Halophila stipulacea in Dominica, West Indies, with a preliminary report from St. Lucia
37447 Journal ArticleWillette, D.A., J. Chalifour, A.O.D. Debrot, M.S. Engel, J. Miller, H.A. Oxenford, F.T. Short, S.C.C. Steiner, F. Védie2014Continued expansion of the trans-Atlantic invasive marine angiosperm Halophila stipulacea in the Eastern Caribbean
45866 Journal ArticleWillette, D.A., K.L. Chiquillo, C. Cross, P. Fong, T. Kelley, C.A. Toline, R. Zweng, and R. Muthukrishnan2020Growth and recovery after small-scale disturbance of a rapidly-expanding invasive seagrass in St. John, U.S. Virgin Islands
46116 Journal ArticleWinters, G., C. Conte, P. Beca-Carretero, H.M. Nguyen, L. Migliore, M. Mulas, G. Rilov, T. Guy-Haim, M.J. González, I. Medina, D. Golomb, N. Baharier, M. Kaminer, and K. Kitson-Walters2023Superior growth traits of invaded (Caribbean) versus native (Red sea) populations of the seagrass Halophila stipulacea


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Citation information: U.S. Geological Survey. [2025]. Nonindigenous Aquatic Species Database. Gainesville, Florida. Accessed [12/17/2025].

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